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<article article-type="research-article" xml:lang="en" dtd-version="1.4"><?da-xref-anchor-style superscripted?><processing-meta base-tagset="archiving" mathml-version="3.0" table-model="xhtml" tagset-family="jats"><restricted-by>pmc</restricted-by></processing-meta><front><journal-meta><journal-id journal-id-type="nlm-ta">Sci Rep</journal-id><journal-id journal-id-type="iso-abbrev">Sci Rep</journal-id><journal-id journal-id-type="pmc-domain-id">1579</journal-id><journal-id journal-id-type="pmc-domain">scirep</journal-id><journal-id journal-id-type="nlm-id">101563288</journal-id><journal-title-group><journal-title>Scientific Reports</journal-title></journal-title-group><issn pub-type="epub">2045-2322</issn><?publisher_abbrev naturepg?><publisher><publisher-name>Nature Publishing Group</publisher-name></publisher></journal-meta><article-meta><article-id pub-id-type="pmcid">PMC11828934</article-id><article-id pub-id-type="pmcid-ver">PMC11828934.1</article-id><article-id pub-id-type="pmcaid">11828934</article-id><article-id pub-id-type="pmcaiid">11828934</article-id><article-id pub-id-type="pmid">39953080</article-id><article-id pub-id-type="doi">10.1038/s41598-025-86974-0</article-id><article-id pub-id-type="publisher-id">86974</article-id><article-version article-version-type="pmc-version">1</article-version><article-categories><subj-group subj-group-type="heading"><subject>Article</subject></subj-group></article-categories><title-group><article-title>Further evidence that averageness and femininity, rather than symmetry and masculinity, predict facial attractiveness judgments</article-title></title-group><contrib-group><contrib contrib-type="author"><name name-style="western"><surname>Lee</surname><given-names initials="P">Pengting</given-names></name><xref ref-type="aff" rid="Aff1"/></contrib><contrib contrib-type="author"><name name-style="western"><surname>Li</surname><given-names initials="J">Jingheng</given-names></name><xref ref-type="aff" rid="Aff1"/></contrib><contrib contrib-type="author"><name name-style="western"><surname>Rafiee</surname><given-names initials="Y">Yasaman</given-names></name><xref ref-type="aff" rid="Aff1"/></contrib><contrib contrib-type="author" corresp="yes"><name name-style="western"><surname>Jones</surname><given-names initials="BC">Benedict C.</given-names></name><address><email>benedict.jones@strath.ac.uk</email></address><xref ref-type="aff" rid="Aff1"/></contrib><contrib contrib-type="author"><name name-style="western"><surname>Shiramizu</surname><given-names initials="VKM">Victor K. M.</given-names></name><xref ref-type="aff" rid="Aff1"/></contrib><aff id="Aff1"><institution-wrap><institution-id institution-id-type="ROR">https://ror.org/00n3w3b69</institution-id><institution-id institution-id-type="GRID">grid.11984.35</institution-id><institution-id institution-id-type="ISNI">0000 0001 2113 8138</institution-id><institution>Department of Psychological Sciences and Health, </institution><institution>University of Strathclyde, </institution></institution-wrap>Glasgow, UK </aff></contrib-group><pub-date pub-type="epub"><day>14</day><month>2</month><year>2025</year></pub-date><pub-date pub-type="collection"><year>2025</year></pub-date><volume>15</volume><issue-id pub-id-type="pmc-issue-id">478255</issue-id><elocation-id>5498</elocation-id><history><date date-type="received"><day>26</day><month>8</month><year>2024</year></date><date date-type="accepted"><day>15</day><month>1</month><year>2025</year></date></history><pub-history><event event-type="pmc-release"><date><day>14</day><month>02</month><year>2025</year></date></event><event event-type="pmc-live"><date><day>15</day><month>02</month><year>2025</year></date></event><event event-type="pmc-last-change"><date iso-8601-date="2025-02-20 02:25:16.283"><day>20</day><month>02</month><year>2025</year></date></event></pub-history><permissions><copyright-statement>© Crown 2025</copyright-statement><copyright-year>2025</copyright-year><license><ali:license_ref xmlns:ali="http://www.niso.org/schemas/ali/1.0/" specific-use="textmining" content-type="ccbylicense">https://creativecommons.org/licenses/by/4.0/</ali:license_ref><license-p><bold>Open Access</bold> This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons licence, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons licence, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons licence and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this licence, visit <ext-link xmlns:xlink="http://www.w3.org/1999/xlink" ext-link-type="uri" xlink:href="https://creativecommons.org/licenses/by/4.0/">http://creativecommons.org/licenses/by/4.0/</ext-link>.</license-p></license></permissions><self-uri xmlns:xlink="http://www.w3.org/1999/xlink" content-type="pmc-pdf" xlink:href="41598_2025_Article_86974.pdf"><?pdf-name 41598_2025_Article_86974.pdf?><?pdf-size 1477216?><?pdf-md5 73315a38be0068877f1c596827583dd8?><?pdf-image-server-status NEVER_LOAD?><?pdf-cloudpmc-urn urn:app:cfd9/11828934/73315a38be00/41598_2025_Article_86974.pdf?></self-uri><abstract id="Abs1"><p id="Par1">Facial attractiveness influences important social outcomes and most studies investigating possible predictors of facial attractiveness have tested for effects of shape symmetry, averageness (i.e., the converse of distinctiveness), and sexual dimorphism (i.e., masculinity–femininity). These studies have typically either tested for these possible effects by experimentally manipulating shape characteristics in faces images or have tested only for bivariate correlations between shape characteristics and attractiveness judgments. However, these two approaches have been criticised for lacking ecological validity and providing little insight into the independent contributions of symmetry, averageness, and sexual dimorphism, respectively. Moreover, the few studies that have investigated the independent contributions of symmetry, averageness, and sexual dimorphism have reported mixed results. Here we measured shape symmetry, averageness, and sexual dimorphism from face images and assessed their independent contribution to attractiveness ratings. Linear mixed effects models showed that facial attractiveness was significantly predicted by averageness in male and female faces and femininity in female faces, but not by masculinity in male faces or symmetry. These results are consistent with other recent work suggesting that averageness and femininity, rather than symmetry and masculinity, predict facial attractiveness.</p></abstract><kwd-group kwd-group-type="npg-subject"><title>Subject terms</title><kwd>Evolution</kwd><kwd>Psychology</kwd></kwd-group><funding-group><award-group><funding-source><institution-wrap><institution-id institution-id-type="FundRef">http://dx.doi.org/10.13039/501100000269</institution-id><institution>Economic and Social Research Council</institution></institution-wrap></funding-source><award-id>ES/X000249/1</award-id></award-group></funding-group><custom-meta-group><custom-meta><meta-name>pmc-status-qastatus</meta-name><meta-value>0</meta-value></custom-meta><custom-meta><meta-name>pmc-status-live</meta-name><meta-value>yes</meta-value></custom-meta><custom-meta><meta-name>pmc-status-embargo</meta-name><meta-value>no</meta-value></custom-meta><custom-meta><meta-name>pmc-status-released</meta-name><meta-value>yes</meta-value></custom-meta><custom-meta><meta-name>pmc-prop-open-access</meta-name><meta-value>yes</meta-value></custom-meta><custom-meta><meta-name>pmc-prop-olf</meta-name><meta-value>no</meta-value></custom-meta><custom-meta><meta-name>pmc-prop-manuscript</meta-name><meta-value>no</meta-value></custom-meta><custom-meta><meta-name>pmc-prop-legally-suppressed</meta-name><meta-value>no</meta-value></custom-meta><custom-meta><meta-name>pmc-prop-has-pdf</meta-name><meta-value>yes</meta-value></custom-meta><custom-meta><meta-name>pmc-prop-has-supplement</meta-name><meta-value>no</meta-value></custom-meta><custom-meta><meta-name>pmc-prop-pdf-only</meta-name><meta-value>no</meta-value></custom-meta><custom-meta><meta-name>pmc-prop-suppress-copyright</meta-name><meta-value>no</meta-value></custom-meta><custom-meta><meta-name>pmc-prop-is-real-version</meta-name><meta-value>no</meta-value></custom-meta><custom-meta><meta-name>pmc-prop-is-scanned-article</meta-name><meta-value>no</meta-value></custom-meta><custom-meta><meta-name>pmc-prop-preprint</meta-name><meta-value>no</meta-value></custom-meta><custom-meta><meta-name>pmc-prop-in-epmc</meta-name><meta-value>yes</meta-value></custom-meta><custom-meta><meta-name>pmc-license-ref</meta-name><meta-value>CC BY</meta-value></custom-meta><custom-meta><meta-name>issue-copyright-statement</meta-name><meta-value>© Springer Nature Limited 2025</meta-value></custom-meta></custom-meta-group></article-meta></front><body><sec id="Sec1"><title>Introduction</title><p id="Par2">Facial-attractiveness judgments influence many important social outcomes<sup><xref ref-type="bibr" rid="CR1">1</xref>–<xref ref-type="bibr" rid="CR3">3</xref></sup>. For example, people prefer to date, mate with, associate with, hire, and vote for individuals with attractive faces<sup><xref ref-type="bibr" rid="CR1">1</xref>–<xref ref-type="bibr" rid="CR3">3</xref></sup>. Given this link between facial attractiveness and social outcomes, many researchers have attempted to identify the physical characteristics that influence facial-attractiveness judgments. Because they are hypothesized to function as cues of immunity to infectious illnesses and/or increase the efficiency with which the perceptual system can encode and process faces (see Rhodes<sup><xref ref-type="bibr" rid="CR3">3</xref></sup> for a discussion of these two accounts of facial-attractiveness judgments), many studies have investigated the possible roles that symmetric, average (i.e., prototypical), and/or sexually dimorphic (i.e., masculine/feminine) shape characteristics play in facial-attractiveness judgments. To elaborate, some researchers have suggested that symmetry, averageness, and sexual dimorphism might influence facial attractiveness judgments because individuals possessing these characteristics also have strong immune systems and, consequently, are produce particularly healthy offspring<sup><xref ref-type="bibr" rid="CR4">4</xref></sup>. However, other researchers have suggested that these characteristics might influence facial attractiveness, not because they advertise aspects of underlying physical condition, but simply because faces possessing these characteristics can be processed most efficiently by the perceptual system<sup><xref ref-type="bibr" rid="CR3">3</xref></sup>.</p><p id="Par3">Many studies have reported that more attractive faces have symmetric<sup><xref ref-type="bibr" rid="CR5">5</xref>–<xref ref-type="bibr" rid="CR8">8</xref></sup> and average<sup><xref ref-type="bibr" rid="CR8">8</xref>–<xref ref-type="bibr" rid="CR10">10</xref></sup> face shapes. Many other studies have reported that more attractive female faces possess feminine face shapes<sup><xref ref-type="bibr" rid="CR11">11</xref>,<xref ref-type="bibr" rid="CR12">12</xref></sup>. By contrast with these results, evidence that more attractive male faces have masculine face shapes is rather mixed (see Little et al.<sup><xref ref-type="bibr" rid="CR2">2</xref></sup> and Rhodes<sup><xref ref-type="bibr" rid="CR3">3</xref></sup> for reviews). For example, while some studies have reported that masculine male faces are perceived to be more attractive than feminine male faces<sup><xref ref-type="bibr" rid="CR11">11</xref></sup>, other studies have reported that feminine male faces are perceived to be more attractive than masculine male faces<sup><xref ref-type="bibr" rid="CR12">12</xref></sup> or that masculinity–femininity does not have a significant effect on male facial attractiveness<sup><xref ref-type="bibr" rid="CR13">13</xref></sup>. However, researchers have recently highlighted two potentially important problems with much of this research on the roles that symmetric, average, and sexually dimorphic shape characteristics play in facial-attractiveness judgments.</p><p id="Par4">First, many studies investigating the roles that symmetric, average, and sexually dimorphic shape characteristics play in facial-attractiveness judgments have done so by experimentally manipulating these characteristics in face images using computer graphics methods<sup><xref ref-type="bibr" rid="CR5">5</xref>,<xref ref-type="bibr" rid="CR9">9</xref>,<xref ref-type="bibr" rid="CR11">11</xref>,<xref ref-type="bibr" rid="CR12">12</xref></sup>. However, researchers have recently raised concerns about the ecological validity of this method<sup><xref ref-type="bibr" rid="CR14">14</xref>–<xref ref-type="bibr" rid="CR21">21</xref></sup>. Indeed, several studies have now demonstrated that the large effects on social judgments of faces that are typically seen when this method is used are considerably smaller, and often not significant, when natural (i.e., unmanipulated) face images are rated and shape characteristics objectively measured from the face images<sup><xref ref-type="bibr" rid="CR14">14</xref>,<xref ref-type="bibr" rid="CR15">15</xref>,<xref ref-type="bibr" rid="CR17">17</xref>,<xref ref-type="bibr" rid="CR22">22</xref>–<xref ref-type="bibr" rid="CR24">24</xref></sup>. This has led some researchers to propose that experimentally manipulating shape characteristics in face images reveals what features participants <italic toggle="yes">can use</italic> to form impressions of others but does not necessarily reveal the features people actually <italic toggle="yes">do use</italic> under more naturalistic viewing condition where faces vary simultaneously on multiple dimensions (for a discussion of this issue, see Satchell et al.<sup><xref ref-type="bibr" rid="CR20">20</xref></sup>).</p><p id="Par5">Second, most studies that have tested for possible relationships between attractiveness ratings of natural (i.e., unmanipulated) faces and measures of shape symmetry, averageness, or sexual dimorphism have done so by testing for bivariate (rather than partial) correlations only (see Rhodes<sup><xref ref-type="bibr" rid="CR3">3</xref></sup> for a review). This approach may be somewhat limited because it is not instructive about which of these face-shape characteristics independently predict attractiveness. Indeed, some studies have reported that averageness and symmetry<sup><xref ref-type="bibr" rid="CR22">22</xref>,<xref ref-type="bibr" rid="CR25">25</xref></sup> and symmetry and sexual dimorphism are positively correlated in face images<sup><xref ref-type="bibr" rid="CR23">23</xref>,<xref ref-type="bibr" rid="CR26">26</xref>–<xref ref-type="bibr" rid="CR28">28</xref></sup>.</p><p id="Par6">Perhaps surprisingly, only a small number of studies have directly addressed these two potentially important issues. However, the specific patterns of results reported differ across these studies. Komori et al.<sup><xref ref-type="bibr" rid="CR29">29</xref></sup> tested for independent relationships between attractiveness ratings of Japanese faces and both averageness and sexual dimorphism, finding that averageness and masculinity (in male faces) and averageness and femininity (in female faces) independently and positively predicted attractiveness ratings. In a follow-up study, Komori et al.<sup><xref ref-type="bibr" rid="CR30">30</xref></sup> reported that averageness and symmetry independently and positively predicted attractiveness ratings of Japanese male faces and that averageness, but not symmetry, independently and positively predicted attractiveness ratings of Japanese female faces. By contrast with these results, Pavlovič et al.<sup><xref ref-type="bibr" rid="CR31">31</xref></sup> tested for independent relationships between attractiveness ratings of Vietnamese faces and symmetry, averageness, and sexual dimorphism, reporting a positive relationship between male attractiveness and averageness, a negative relationship between male attractiveness and masculinity, and a positive relationship between female attractiveness and averageness. By contrast, femininity was not significantly correlated with female attractiveness and symmetry was not significantly correlated with facial attractiveness in either sex. However, a different pattern of results was reported by Kleisner et al.<sup><xref ref-type="bibr" rid="CR32">32</xref></sup> in a study using a set of faces of diverse ethnicities. In this study, Kleisner et al.<sup><xref ref-type="bibr" rid="CR32">32</xref></sup> reported independent positive relationships between female attractiveness and averageness and femininity, but not symmetry, and independent positive relationships between male attractiveness and averageness, but not masculinity or symmetry. Thus, while each of these studies reported that averageness was positively correlated with facial attractiveness, findings for symmetry and both masculinity (in male faces) and femininity (in female faces) were mixed across studies.</p><p id="Par7">In light of the mixed results described above, the current study tested for independent relationships between attractiveness ratings of white male and female faces and objective measures of asymmetry, distinctiveness (the converse of averageness), and sexual dimorphism (i.e., masculinity/femininity) of face shape. Given the mixed results from previous studies, we made no specific predictions about which particular features will or will not predict attractiveness significantly in this exploratory study.</p></sec><sec id="Sec2"><title>Methods</title><sec id="Sec3"><title>Stimuli</title><p id="Par8">Stimuli were face images of 50 white men (mean age = 24.2 years, SD = 3.99 years) and 50 white women (mean age = 24.3 years, SD = 4.01 years) that were obtained from an open-access face-image database<sup><xref ref-type="bibr" rid="CR33">33</xref></sup>. Individuals posed front-on to the camera, with direct gaze, and with neutral expressions. Images were standardized on pupil position and clothing was masked prior to rating. Example images are shown in Fig. <xref rid="Fig1" ref-type="fig">1</xref>. Images (and the shape templates we used to calculate masculinity, distinctiveness, and asymmetry scores) are publicly available at <ext-link xmlns:xlink="http://www.w3.org/1999/xlink" ext-link-type="uri" xlink:href="https://osf.io/a3947/">https://osf.io/a3947/</ext-link>.<fig id="Fig1" position="float" orientation="portrait"><label>Fig. 1</label><caption><p>Examples of male (top row) and female (bottom row) face images used in our study.</p></caption><graphic xmlns:xlink="http://www.w3.org/1999/xlink" id="d33e349" position="float" orientation="portrait" xlink:href="41598_2025_86974_Fig1_HTML.jpg"><?image-name 41598_2025_86974_Fig1_HTML.jpg?><?image-size 158663?><?image-md5 8c063616999ae1d3eb95dc21414bdc67?><?image-image-server-status LOAD_COMPLETED?><?image-original-height 1729?><?image-original-width 1182?><?image-scaled-height 1153?><?image-scaled-width 788?><?image-cloudpmc-urn urn:cdn:blobs/cfd9/11828934/8c063616999a/41598_2025_86974_Fig1_HTML.jpg?><?thumb-name 41598_2025_86974_Fig1_HTML.gif?><?thumb-size 8976?><?thumb-md5 8ed40649170a4df8dc294c6f74e7ccc8?><?thumb-image-server-status NEVER_LOAD?><?thumb-scaled-height 146?><?thumb-scaled-width 100?><?thumb-cloudpmc-urn urn:cdn:blobs/cfd9/11828934/8ed40649170a/41598_2025_86974_Fig1_HTML.gif?></graphic></fig></p></sec><sec id="Sec4"><title>Attractiveness ratings</title><p id="Par9">Two hundred heterosexual men and 200 heterosexual women (mean age = 24.83 years, SD = 5.70 years) were randomly allocated to rate either the 50 male or 50 female face images in an online study using a 7-point scale (1 = much less attractive than average, 7 = much more attractive than average). Trial order was fully randomised. Cronbach’s alphas were high for ratings of both male (Cronbach’s alpha = 0.99) and female (Cronbach’s alpha = 0.99) face images. Cronbach’s alphas were also high for each combination of face sex and rater sex when calculated separately (all Cronbach’s alphas &gt; 0.97). The mean attractiveness rating for male faces was 2.64 (SD = 1.46) and the mean attractiveness rating for female faces was 2.71 (SD = 1.48). Attractiveness ratings were collected using the Experimentum data-collection platform<sup><xref ref-type="bibr" rid="CR34">34</xref></sup>.</p></sec><sec id="Sec5"><title>Measuring face-shape sexual dimorphism</title><p id="Par10">Sexual dimorphism of face shape was objectively measured for each of the 50 male and 50 female face images using the facefuns package<sup><xref ref-type="bibr" rid="CR35">35</xref></sup> in R<sup><xref ref-type="bibr" rid="CR36">36</xref></sup>. This method has been used in many previous studies to assess sexual dimorphism of face shape<sup><xref ref-type="bibr" rid="CR14">14</xref>,<xref ref-type="bibr" rid="CR22">22</xref>,<xref ref-type="bibr" rid="CR23">23</xref>,<xref ref-type="bibr" rid="CR37">37</xref>–<xref ref-type="bibr" rid="CR39">39</xref></sup>. Shape components were first derived from Principal Component Analysis (PCA) of 132 Procrustes-aligned landmark points (see Holzleitner et al.<sup><xref ref-type="bibr" rid="CR38">38</xref></sup> for a diagram showing these facial landmarks) on each of the 50 male and 50 female face images. Scores representing sexual dimorphism of face shape were then calculated from each photograph using a vector analysis method<sup><xref ref-type="bibr" rid="CR14">14</xref>,<xref ref-type="bibr" rid="CR22">22</xref>,<xref ref-type="bibr" rid="CR23">23</xref>,<xref ref-type="bibr" rid="CR37">37</xref>–<xref ref-type="bibr" rid="CR39">39</xref></sup>. This method uses the shape principal components to locate each face on a female-male continuum. The female-male continuum was defined by calculating the average shape information of the 50 female faces and the average shape information of the 50 male faces. Sexual dimorphism scores were then derived by projecting each image onto this female-male vector. Higher scores indicated more masculine face shapes. No scores were more than three standard deviations from the mean (i.e., there were no extreme values).</p></sec><sec id="Sec6"><title>Measuring face-shape distinctiveness</title><p id="Par11">Distinctiveness scores were also calculated from each photograph using the facefuns package<sup><xref ref-type="bibr" rid="CR35">35</xref></sup> in R<sup><xref ref-type="bibr" rid="CR36">36</xref></sup>. This technique has been used to measure face-shape distinctiveness in many previous studies<sup><xref ref-type="bibr" rid="CR22">22</xref>,<xref ref-type="bibr" rid="CR23">23</xref>,<xref ref-type="bibr" rid="CR37">37</xref>,<xref ref-type="bibr" rid="CR38">38</xref></sup>. This method uses the shape principal components described in the previous section of our methods to measure the distance each face lies from the mathematical average shape for the sample of faces of the same sex. That is, the average shape values for the same-sex sample were calculated and, for each image, the Euclidean distance from the average was derived. Higher scores indicate that the face lies a further distance away from the average (i.e., had a more distinctive shape). We measured distinctiveness scores for male and female faces separately in light of evidence that faces are primarily processed relative to sex-specific prototypes<sup><xref ref-type="bibr" rid="CR40">40</xref>,<xref ref-type="bibr" rid="CR41">41</xref></sup>. No scores were more than three standard deviations from the mean (i.e., there were no extreme values).</p></sec><sec id="Sec7"><title>Measuring face-shape asymmetry</title><p id="Par12">Asymmetry scores were also calculated from each photograph using the facefuns package<sup><xref ref-type="bibr" rid="CR35">35</xref></sup> in R<sup><xref ref-type="bibr" rid="CR36">36</xref></sup>. This technique has been used to measure face-shape asymmetry in many previous studies<sup><xref ref-type="bibr" rid="CR22">22</xref>,<xref ref-type="bibr" rid="CR37">37</xref>,<xref ref-type="bibr" rid="CR38">38</xref></sup>. For each image, the landmark template was mirrored, and shape asymmetry measured as the Euclidean distance between original and mirrored templates. Higher scores indicate that the face has greater asymmetry. One extreme value (i.e., one score more than three standard deviations from the mean) was adjusted (i.e., winsorized) to be three standard deviations from the mean prior to further analyses.</p></sec></sec><sec id="Sec8"><title>Results</title><p id="Par13">All statistical analyses were carried out using R<sup><xref ref-type="bibr" rid="CR36">36</xref></sup>, with the packages lme4<sup><xref ref-type="bibr" rid="CR42">42</xref></sup>, lmerTest 3.1-3<sup><xref ref-type="bibr" rid="CR43">43</xref></sup>, jtools 2.2.3<sup><xref ref-type="bibr" rid="CR44">44</xref></sup>, and robustHD 0.7.3<sup><xref ref-type="bibr" rid="CR45">45</xref></sup>. Data processing and display used kableExtra 1.3.4<sup><xref ref-type="bibr" rid="CR46">46</xref></sup> and tidyverse 1.3.1<sup><xref ref-type="bibr" rid="CR47">47</xref></sup>. All data, full outputs, and analysis code are publicly available on the Open Science Framework (<ext-link xmlns:xlink="http://www.w3.org/1999/xlink" ext-link-type="uri" xlink:href="https://osf.io/xuz9r/">https://osf.io/xuz9r/</ext-link>).</p><p id="Par14">Attractiveness ratings served as the dependent variable in a linear mixed effects model. The model included main effects of sexual dimorphism scores, distinctiveness scores, asymmetry scores, rater sex (effect coded so that − 0.5 corresponded to male raters and 0.5 corresponded to female raters), and face sex (effect coded so that − 0.5 corresponded to male faces and 0.5 corresponded to female faces) as predictors, as well as all possible two- and three-way interactions (excluding those involving multiple continuous predictors). The model also included by-rater and by-stimulus random intercepts, by-rater random slopes for sexual dimorphism, asymmetry, and distinctiveness (face sex varied between raters), and by-stimulus random slopes for rater sex. Sexual dimorphism, distinctiveness scores, and asymmetry scores were standardised prior to analyses by converting them to z scores (separately for each face sex). Full results for this model are summarised in Table <xref rid="Tab1" ref-type="table">1</xref>. This analysis revealed significant negative main effects of both distinctiveness and sexual dimorphism. By contrast, the effect of asymmetry was not significant. The only significant interaction was the two-way interaction between sexual dimorphism and face sex. The significant effect of distinctiveness is shown in Fig. <xref rid="Fig2" ref-type="fig">2</xref>.<table-wrap id="Tab1" position="float" orientation="portrait"><label>Table 1</label><caption><p>Results of our analysis of attractiveness ratings.</p></caption><table frame="hsides" rules="groups"><thead><tr><th align="left" colspan="1" rowspan="1"/><th align="left" colspan="1" rowspan="1">Estimate</th><th align="left" colspan="1" rowspan="1">Standard error</th><th align="left" colspan="1" rowspan="1">t value</th><th align="left" colspan="1" rowspan="1">df</th><th align="left" colspan="1" rowspan="1"><italic toggle="yes">p</italic> value</th></tr></thead><tbody><tr><td align="left" colspan="1" rowspan="1">Intercept</td><td char="." align="char" colspan="1" rowspan="1">2.466</td><td char="." align="char" colspan="1" rowspan="1">0.108</td><td char="." align="char" colspan="1" rowspan="1">22.766</td><td char="." align="char" colspan="1" rowspan="1">131.071</td><td char="." align="char" colspan="1" rowspan="1">&lt; 0.001</td></tr><tr><td align="left" colspan="1" rowspan="1">Distinctiveness</td><td char="." align="char" colspan="1" rowspan="1">− 0.255</td><td char="." align="char" colspan="1" rowspan="1">0.062</td><td char="." align="char" colspan="1" rowspan="1">− 4.099</td><td char="." align="char" colspan="1" rowspan="1">100.778</td><td char="." align="char" colspan="1" rowspan="1">&lt; 0.001</td></tr><tr><td align="left" colspan="1" rowspan="1">Face sex</td><td char="." align="char" colspan="1" rowspan="1">− 0.479</td><td char="." align="char" colspan="1" rowspan="1">0.217</td><td char="." align="char" colspan="1" rowspan="1">− 2.212</td><td char="." align="char" colspan="1" rowspan="1">131.071</td><td char="." align="char" colspan="1" rowspan="1">0.029</td></tr><tr><td align="left" colspan="1" rowspan="1">Rater sex</td><td char="." align="char" colspan="1" rowspan="1">− 0.188</td><td char="." align="char" colspan="1" rowspan="1">0.085</td><td char="." align="char" colspan="1" rowspan="1">− 2.227</td><td char="." align="char" colspan="1" rowspan="1">428.086</td><td char="." align="char" colspan="1" rowspan="1">0.026</td></tr><tr><td align="left" colspan="1" rowspan="1">Sexual dimorphism</td><td char="." align="char" colspan="1" rowspan="1">− 0.278</td><td char="." align="char" colspan="1" rowspan="1">0.100</td><td char="." align="char" colspan="1" rowspan="1">− 2.783</td><td char="." align="char" colspan="1" rowspan="1">101.241</td><td char="." align="char" colspan="1" rowspan="1">0.006</td></tr><tr><td align="left" colspan="1" rowspan="1">Asymmetry</td><td char="." align="char" colspan="1" rowspan="1">− 0.092</td><td char="." align="char" colspan="1" rowspan="1">0.068</td><td char="." align="char" colspan="1" rowspan="1">− 1.341</td><td char="." align="char" colspan="1" rowspan="1">100.493</td><td char="." align="char" colspan="1" rowspan="1">0.183</td></tr><tr><td align="left" colspan="1" rowspan="1">Distinctiveness × face sex</td><td char="." align="char" colspan="1" rowspan="1">0.118</td><td char="." align="char" colspan="1" rowspan="1">0.124</td><td char="." align="char" colspan="1" rowspan="1">0.949</td><td char="." align="char" colspan="1" rowspan="1">100.778</td><td char="." align="char" colspan="1" rowspan="1">0.345</td></tr><tr><td align="left" colspan="1" rowspan="1">Distinctiveness × rater sex</td><td char="." align="char" colspan="1" rowspan="1">− 0.023</td><td char="." align="char" colspan="1" rowspan="1">0.022</td><td char="." align="char" colspan="1" rowspan="1">− 1.083</td><td char="." align="char" colspan="1" rowspan="1">125.376</td><td char="." align="char" colspan="1" rowspan="1">0.281</td></tr><tr><td align="left" colspan="1" rowspan="1">Face sex × rater sex</td><td char="." align="char" colspan="1" rowspan="1">0.239</td><td char="." align="char" colspan="1" rowspan="1">0.169</td><td char="." align="char" colspan="1" rowspan="1">1.411</td><td char="." align="char" colspan="1" rowspan="1">428.086</td><td char="." align="char" colspan="1" rowspan="1">0.159</td></tr><tr><td align="left" colspan="1" rowspan="1">Sexual dimorphism × face sex</td><td char="." align="char" colspan="1" rowspan="1">− 0.53</td><td char="." align="char" colspan="1" rowspan="1">0.200</td><td char="." align="char" colspan="1" rowspan="1">− 2.653</td><td char="." align="char" colspan="1" rowspan="1">101.241</td><td char="." align="char" colspan="1" rowspan="1">0.009</td></tr><tr><td align="left" colspan="1" rowspan="1">Sexual dimorphism × rater sex</td><td char="." align="char" colspan="1" rowspan="1">0.035</td><td char="." align="char" colspan="1" rowspan="1">0.036</td><td char="." align="char" colspan="1" rowspan="1">0.970</td><td char="." align="char" colspan="1" rowspan="1">134.608</td><td char="." align="char" colspan="1" rowspan="1">0.334</td></tr><tr><td align="left" colspan="1" rowspan="1">Asymmetry × face sex</td><td char="." align="char" colspan="1" rowspan="1">0.03</td><td char="." align="char" colspan="1" rowspan="1">0.137</td><td char="." align="char" colspan="1" rowspan="1">0.222</td><td char="." align="char" colspan="1" rowspan="1">100.493</td><td char="." align="char" colspan="1" rowspan="1">0.825</td></tr><tr><td align="left" colspan="1" rowspan="1">Asymmetry × rater sex</td><td char="." align="char" colspan="1" rowspan="1">− 0.007</td><td char="." align="char" colspan="1" rowspan="1">0.023</td><td char="." align="char" colspan="1" rowspan="1">− 0.320</td><td char="." align="char" colspan="1" rowspan="1">117.682</td><td char="." align="char" colspan="1" rowspan="1">0.750</td></tr><tr><td align="left" colspan="1" rowspan="1">Distinctiveness × face sex × rater sex</td><td char="." align="char" colspan="1" rowspan="1">0.001</td><td char="." align="char" colspan="1" rowspan="1">0.043</td><td char="." align="char" colspan="1" rowspan="1">− 0.010</td><td char="." align="char" colspan="1" rowspan="1">125.376</td><td char="." align="char" colspan="1" rowspan="1">0.992</td></tr><tr><td align="left" colspan="1" rowspan="1">Sexual dimorphism × face sex × rater sex</td><td char="." align="char" colspan="1" rowspan="1">0.055</td><td char="." align="char" colspan="1" rowspan="1">0.072</td><td char="." align="char" colspan="1" rowspan="1">0.766</td><td char="." align="char" colspan="1" rowspan="1">134.608</td><td char="." align="char" colspan="1" rowspan="1">0.445</td></tr><tr><td align="left" colspan="1" rowspan="1">Asymmetry × face sex × rater sex</td><td char="." align="char" colspan="1" rowspan="1">0.037</td><td char="." align="char" colspan="1" rowspan="1">0.047</td><td char="." align="char" colspan="1" rowspan="1">0.797</td><td char="." align="char" colspan="1" rowspan="1">117.682</td><td char="." align="char" colspan="1" rowspan="1">0.427</td></tr></tbody></table></table-wrap><fig id="Fig2" position="float" orientation="portrait"><label>Fig. 2</label><caption><p>The significant negative effect of face-shape distinctiveness on attractiveness. The shaded area shows the 95% confidence interval.</p></caption><graphic xmlns:xlink="http://www.w3.org/1999/xlink" id="d33e749" position="float" orientation="portrait" xlink:href="41598_2025_86974_Fig2_HTML.jpg"><?image-name 41598_2025_86974_Fig2_HTML.jpg?><?image-size 37090?><?image-md5 13e7013ec1d512c8fa84fbeba577e9fc?><?image-image-server-status LOAD_COMPLETED?><?image-original-height 833?><?image-original-width 1182?><?image-scaled-height 555?><?image-scaled-width 788?><?image-cloudpmc-urn urn:cdn:blobs/cfd9/11828934/13e7013ec1d5/41598_2025_86974_Fig2_HTML.jpg?><?thumb-name 41598_2025_86974_Fig2_HTML.gif?><?thumb-size 2012?><?thumb-md5 1de3f0cc9e2f5a562f431cbbd35c73fd?><?thumb-image-server-status NEVER_LOAD?><?thumb-scaled-height 80?><?thumb-scaled-width 113?><?thumb-cloudpmc-urn urn:cdn:blobs/cfd9/11828934/1de3f0cc9e2f/41598_2025_86974_Fig2_HTML.gif?></graphic></fig></p><p id="Par15">To interpret the significant interaction between sexual dimorphism and face sex, we repeated the analysis described above, this time analysing male and female faces in separate models. For female faces, there was a significant negative effect of sexual dimorphism (estimate = − 0.543, SE = 0.143, t = − 3.803, df = 50.802, <italic toggle="yes">p</italic> &lt; 0.001). By contrast, the effect of sexual dimorphism was not significant for male faces (estimate = − 0.013, SE = 0.138, t = − 0.094, df = 50.145, <italic toggle="yes">p</italic> = 0.926). These results are shown in Fig. <xref rid="Fig3" ref-type="fig">3</xref>. Full results for both models are given on the Open Science Framework (<ext-link xmlns:xlink="http://www.w3.org/1999/xlink" ext-link-type="uri" xlink:href="https://osf.io/xuz9r/">https://osf.io/xuz9r/</ext-link>).<fig id="Fig3" position="float" orientation="portrait"><label>Fig. 3</label><caption><p>The significant effect of face-shape sexual dimorphism on female attractiveness and non-significant effect of face-shape sexual dimorphism on male attractiveness. The shaded areas show the 95% confidence intervals.</p></caption><graphic xmlns:xlink="http://www.w3.org/1999/xlink" id="d33e771" position="float" orientation="portrait" xlink:href="41598_2025_86974_Fig3_HTML.jpg"><?image-name 41598_2025_86974_Fig3_HTML.jpg?><?image-size 45691?><?image-md5 8e5e98b18d9c12828639df67fd202ee9?><?image-image-server-status LOAD_COMPLETED?><?image-original-height 848?><?image-original-width 1182?><?image-scaled-height 565?><?image-scaled-width 788?><?image-cloudpmc-urn urn:cdn:blobs/cfd9/11828934/8e5e98b18d9c/41598_2025_86974_Fig3_HTML.jpg?><?thumb-name 41598_2025_86974_Fig3_HTML.gif?><?thumb-size 2159?><?thumb-md5 8c943b3d5caee595839c25229c0c4532?><?thumb-image-server-status NEVER_LOAD?><?thumb-scaled-height 80?><?thumb-scaled-width 111?><?thumb-cloudpmc-urn urn:cdn:blobs/cfd9/11828934/8c943b3d5cae/41598_2025_86974_Fig3_HTML.gif?></graphic></fig></p><p id="Par16">We also repeated the initial analysis described above, this time including all possible interactions among predictors in the model. This analysis revealed significant interactions among distinctiveness, rater sex, and asymmetry and among distinctiveness, face sex, rater sex, and sexual dimorphism. Results of these analyses are reported in full at <ext-link xmlns:xlink="http://www.w3.org/1999/xlink" ext-link-type="uri" xlink:href="https://osf.io/xuz9r/">https://osf.io/xuz9r/</ext-link>. Because these high-order interactions were unexpected, we do not discuss them further.</p></sec><sec id="Sec9"><title>Discussion</title><p id="Par17">Our analyses of facial-attractiveness ratings showed a significant positive relationship between averageness of face shape and attractiveness ratings of male and female faces (i.e., negative relationships between distinctiveness and facial attractiveness) and a significant positive effect of femininity of face shape and attractiveness ratings of female faces. By contrast, masculinity of face shape was not significantly correlated with male facial attractiveness and symmetry was not significantly correlated with either male or female attractiveness. This pattern of results (significant effects of averageness and female femininity, but not symmetry or male masculinity) is identical to those reported previously by Kleisner et al.<sup><xref ref-type="bibr" rid="CR32">32</xref></sup>. Although other studies testing for independent effects of averageness, symmetry, and sexual dimorphism of face shape on attractiveness ratings also reported positive effects of averageness<sup><xref ref-type="bibr" rid="CR29">29</xref>–<xref ref-type="bibr" rid="CR31">31</xref></sup> and female femininity<sup><xref ref-type="bibr" rid="CR29">29</xref></sup>, we did not observe the significant effects of symmetry<sup><xref ref-type="bibr" rid="CR30">30</xref></sup> or male masculinity<sup><xref ref-type="bibr" rid="CR29">29</xref>,<xref ref-type="bibr" rid="CR31">31</xref></sup> that these studies reported. Thus, our findings add to a growing body of research suggesting that averageness is a key shape characteristic for facial attractiveness judgments and that there is little evidence for a significant role of symmetry or male masculinity (see also Kleisner et al.<sup><xref ref-type="bibr" rid="CR32">32</xref></sup>).</p><p id="Par18">There has been considerable debate in the facial-attractiveness literature regarding the extent to which preferences for faces with average shapes are due to a hypothesized positive effect of symmetry on attractiveness ratings<sup><xref ref-type="bibr" rid="CR48">48</xref>–<xref ref-type="bibr" rid="CR50">50</xref></sup>. However, results from studies in which averageness was experimentally manipulated independently of symmetry suggest that symmetry contributes little to the positive effect of averageness on facial-attractiveness judgments<sup><xref ref-type="bibr" rid="CR48">48</xref>–<xref ref-type="bibr" rid="CR50">50</xref></sup>. Our finding that averageness, but not symmetry, has an independent positive effect on attractiveness ratings provides further evidence that averageness is attractive independent of the putative effects of symmetry and suggests that symmetry may, in fact, contribute little to the positive effect of averageness on facial attractiveness (see also Kleisner et al.<sup><xref ref-type="bibr" rid="CR32">32</xref></sup>). Although some previous studies have reported that symmetry influences facial attractiveness judgments (see Rhodes<sup><xref ref-type="bibr" rid="CR3">3</xref></sup> for a meta-analytic review), those studies did not typically control for the positive effects of averageness on attractiveness judgments. This issue is potentially noteworthy, since averageness and symmetry in faces are correlated<sup><xref ref-type="bibr" rid="CR22">22</xref>,<xref ref-type="bibr" rid="CR25">25</xref></sup>. Indeed, recent work has suggested that effects of averageness mediate effects of symmetry on social judgments of faces<sup><xref ref-type="bibr" rid="CR23">23</xref></sup>. Whether this mediation is due to symmetry and averageness having similar effects on the ease with which faces can be processed by the perceptual system or that both characteristics are associated with similar aspects of physical condition remains an open question. Similarly, whether averageness influences facial attractiveness because average faces can be processed more efficiently or because it advertises physical condition (the two most common explanations for effects of averageness on facial attractiveness, Rhodes<sup><xref ref-type="bibr" rid="CR3">3</xref></sup>) is likely to be an important topic for future research. This issue is also an important direction for future research on the effects of femininity on women’s facial attractiveness.</p><p id="Par19">Our null result for masculinity and male facial attractiveness contrasts with findings from some previous research that found male facial attractiveness was influenced by masculine shape characteristics<sup><xref ref-type="bibr" rid="CR11">11</xref>,<xref ref-type="bibr" rid="CR12">12</xref>,<xref ref-type="bibr" rid="CR51">51</xref></sup>. However, studies showing this latter pattern of results typically employed stimuli in which shape characteristics were experimentally manipulated. Several lines of evidence suggest that findings obtained using this type of manipulated stimuli generalise poorly to studies investigating judgments of natural (i.e., unmanipulated) stimuli, as were employed in the current work (see Dong et al.<sup><xref ref-type="bibr" rid="CR14">14</xref></sup> for a recent review of work on this issue). Nonetheless, while we suggest that differences between the results of the current study and those of previous research on preferences for male facial masculinity may be due to this methodological issue, we also acknowledge that putative effects of contextual factors (e.g., sociocultural factors) on preferences for male facial masculinity<sup><xref ref-type="bibr" rid="CR52">52</xref></sup> might also contribute to differences in findings across studies.</p><p id="Par20">Although we observed significant relationships between facial attractiveness and face-shape averageness and between shape femininity and female facial attractiveness, these effects were quite weak (see also Kleisner et al.<sup><xref ref-type="bibr" rid="CR32">32</xref></sup>). This issue raises the question of what additional factors might contribute to facial-attractiveness judgments. While most research on facial attractiveness has focussed on investigating possible roles of averageness, sexual dimorphism, and symmetry of face shape, other studies have suggested that surface information (e.g., color and texture information in faces) and cues of adiposity (i.e., facial correlates of body mass index) both may influence facial-attractiveness judgments (see Holzleitner et al.<sup><xref ref-type="bibr" rid="CR38">38</xref></sup> for a review). Considering these (and other) additional characteristics may improve the predictive power of models of facial attractiveness.</p><p id="Par21">To summarise, much of the research on predictors of facial-attractiveness judgments has focussed on the possible roles played by symmetry, averageness, and sexual dimorphism in shape characteristics. However, relatively few studies have investigated the extent to which these face-shape characteristics independently predict attractiveness and those studies that have done so have reported somewhat mixed results<sup><xref ref-type="bibr" rid="CR29">29</xref>–<xref ref-type="bibr" rid="CR32">32</xref></sup>. Our finding that facial attractiveness is significantly correlated with averageness in both male and female faces and femininity in female faces, but not masculinity in male faces or symmetry directly replicates the pattern of results reported by Kleisner et al.<sup><xref ref-type="bibr" rid="CR32">32</xref></sup>. Thus, our results present further evidence that averageness and femininity, rather than symmetry and masculinity, predict facial-attractiveness judgments.</p></sec></body><back><fn-group><fn><p><bold>Publisher’s note</bold></p><p>Springer Nature remains neutral with regard to jurisdictional claims in published maps and institutional affiliations.</p></fn></fn-group><ack><title>Acknowledgements</title><p>This research was supported by ESRC grant ES/X000249/1 awarded to BCJ. For the purpose of Open Access, the authors have applied a Creative Commons Attribution (CC BY) to any Author Accepted Manuscript (AAM) version arising from this submission.</p></ack><notes notes-type="author-contribution"><title>Author contributions</title><p>PL, VS, and BJ designed the study, with feedback from YR and JL. VS, PL, and YR carried out analyses. PL and BJ wrote the main manuscript. 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